Karen Beard

Publications | Book Chapters

• Beard, K.H, (2024). An out-of-towner settles in: Coqui frogs from Puerto Rico find paradise in Hawaii: Amphibian Biology, Volume 9 (Status of Conservation and Decline of Amphibians in the Western Hemisphere), Part 7 (Canada and the United States) of the Amphibian Biology World Encyclopedia. Olson, D.H. and D.S. Pilliod, Amphibians of the US West: Biogeography, Biodiversity, Threats, and Conservation
• Beard, K.H, (2017). Chapter 9: Frogs (Coqui Frogs, Greenhouse Frogs, Cuban Tree Frogs, and Cane Toads): Ecology and Management of Terrestrial Vertebrate Invasive Species in the United States . CRC Press/Taylor & Francis Group
• Edwards, T.C, Cutler, D., Beard, K.H, (2014). Predicting Risk of Invasive Species Occurrence: Plot Based: Early detection of invasive plants—Principles and practices. USGS/NPS joint publication
• Edwards, T.C, Cutler, D., Beard, K.H, (2014). The Process of Model Assessment and Evaluation: Early detection of invasive plants—Principles and practices.
• Beard, K.H, Pitt, W.C, (2012). Chapter 26: Caribbean tree frog (Eleutherodactylus coqui): Handbook of Global Freshwater Invasive Species. Earthscan, London
• Scatena, F.N, Blanco, J.F, Beard, K.H, Waide, R., Lugo, A.E, Brokaw, N., Silver, W., Haines, B., ZImmerman, J., (2012). Chapter 4: Disturbance regime: A Caribbean Forest Tapestry: The Multidimensional Nature of Disturbance and Response. Oxford University Press
• Crowl, T.A, Brokaw, N., Waide, R., Gonzalez, G., Beard, K.H, Greathouse, E., Lugo, A.E, Covich, A.P, Lodge, J., Pringle, C., Thompson, J., Belovsky, G.E, Brokaw, J.N, Crowl, T., Lugo, A., McDowell, W.M, Scatena, F., Waide, R.E, Willig, M.R, (2012). Chapter 6: When and where biota matter: linking disturbance regimes, species characteristics and dynamics of communities and ecosystems: A Caribbean Forest Tapestry: The Multidimensional Nature of Disturbance and Response. Oxford University Press
• Edwards, T.C, Cutler, D., Beard, K.H, Gibson, J., (2010). Predicting Invasive Plant Species in the Klamath Network National Parks: A Process for Prioritizing Curriculum Vitae for D. Richard Cutler November 2010 6 Prevention: Early Detection of Invasive Plants Handbook. USGS/NPS joint publication
• Beard, K.H, (2006). Case Study Box: Puerto Rico and Hawaii: Wet tropical forests and the dilemma of coqui frog conservation and eradication: Forests and Society: Sustainability and life cycles of forests in human landscapes. *
• Vogt, K.A, Schmitz, O., Beard, K.H, O'Hara, J.L, Booth, M., (2001). Conservation Biology, Contemporary: Encyclopedia of Biodiversity. Encyclopedia of Biodiversity
• Beard, K.H, (1999). Keystone species, ecosystem engineers: where single species are good indicators of the ecosystem. In: Forest Certification: Roots, Issues, Challenges and Benefits (K.A. Vogt, B.C Larson, et al., Eds.)

An asterisk (*) at the end of a publication indicates that it has not been peer-reviewed.

Publications | Journal Articles

Academic Journal

• Garrett, S., Beard, K.H, Du Toit, J.T, (2025). Quantification of the effect of lagomorph herbivory on grazing resources in a semi-arid rangeland. Ecosphere, 16:11, e70463. doi: DOI: 10.1002/ecs2.70463
• Ohlert, T., Smith, M.D, Wilkins, K.D, Holdrege, M.C, Wilfahrt, P., Collins, S.L, Knapp, A.K, Sala, O.E, Dukes, J.S, Phillips, R.P, Yahdjian, L., Gherardi, L.A, Beier, C., Fraser, L.H, Jentsch, A., Loik, M.E, Maestre, F.T, Power, S.A, Yu, Q., Felton, A.J, Munson, S.M, Luo, Y., Abdoli, H., Abedi, M., Alados, C.L, Alberti, J., Alon, M., An, H., Anacker, B., Anderson, M., Auge, H., Bachle, S., Bahalkeh, K., Bahn, M., Batbaatar, A., Bauerle, T., Beard, K.H, Behn, K., Beil, I., Biancari, L., Blindow, I., Bondaruk, V.F, Borer, E.T, Bork, E.W, Bruschetti, C.M, Byrne, K.M, Cahill, Jr., J.F, Calvo, D.A, Carbognani, M., Cardoni, A., Carlyle, C.N, Castillo-Garcia, M., Chang, S.X, Chieppa, J., Cianciaruso, M.V, Cohen, O., Cordeiro, A.L, Cusack, D.F, Dahlke, S., Daleo, P., D'Antonio, C.M, Dietterich, L.H, Doherty, T.S, Dubbert, M., Ebeling, A., Eisenhauer, N., Fischer, F.M, Forte, T.G, Gebauer, T., Gozalo, B., Greenville, A.C, Guidoni-Martins, K.G, Hannusch, H.J, Haugum, S.V, Hautier, Y., Hefting, M., Henry, H.A, Hoss, D., Ingrisch, J., Iribarne, O., Isbell, F., Johnson, Y., Jordan, S., Kelly, E.F, Kimmel, K., Kreyling, J., Kröel-Dulay, G., Kröpfl, A., Kübert, A., Kulmatiski, A., Lamb, E.G, Larsen, K.S, Larson, J., Lawson, J., Leder, C.V, Linstädter, A., Liu, J., Liu, S., Lodge, A.G, Longo, G., Loydi, A., Luan, J., Lubbe, F.C, Macfarlane, C., Mackie-Haas, K., Malyshev, A.V, Maturano-Ruiz, A., Merchant, T., Metcalfe, D.B, Mori, A.S, Mudongo, E., Newman, G.S, Nielsen, U.N, Nimmo, D., Niu, Y., Nobre, P., O'Connor, R.C, Ogaya, R., Oñatibia, G.R, Orbán, I., Osborne, B., Otfinowski, R., Pärtel, M., Penuelas, J., Peri, P.L, Peter, G., Petraglia, A., Picon-Cochard, C., Pillar, V.D, Piñeiro-Guerra, J.M, Ploughe, L.W, Plowes, R.M, Portales-Reyes, C., Prober, S.M, Pueyo, Y., Reed, S.C, Ritchie, E.G, Rodríguez, D.A, Rogers, W.E, Roscher, C., Sánchez, A.M, Santos, B.A, Scarfó, M.C, Seabloom, E.W, Shi, B., Souza, L., Stampfli, A., Standish, R.J, Sternberg, M., Sun, W., Sünnemann, M., Tedder, M., Thorvaldsen, P., Tian, D., Tielbörger, K., Valdecantos, A., Liesbeth van den Brink, , Vandvik, V., Vankoughnett, M.R, Velle, L.G, Wang, C., Wang, Y., Wardle, G.M, Werner, C., Wei, C., Wiehl, G., Williams, J.L, Wolf, A.A, Zeiter, M., Zhang, F., Zhu, J., Zong, N., Zuo, X., (2025). Drought intensity and duration interact to magnify losses in primary productivity. Science, 390:6770, 284-289.
• Kelsey, K.C, Leffler, A.J, Petit Bon, M., Barr, B.N, Beard, K.H, (2025). Responding to rising waters and temperatures: Greenhouse gas flux from a high-latitude coastal wetland. Environmental Research Letter, 20:10, 104040. doi: https://doi.org/10.1088/1748-9326/adff98
• Barr, B.N, Kelsey, K.C, Leffler, A.J, Petit Bon, M., Beard, K.H, (2025). Salinity and moisture influence CO2 and CH4 emissions from high latitude coastal soils in high-latitude coastal wetland. JGR Biogeosciences, 130:7, e2024JG008629. doi: https://doi.org/10.1029/2024JG008629
• Chavez, E.A, Adkins, J., Waring, B., Beard, K.H, Miller, L., Choi, R., Saunders, T., Atwood, T., (2025). Herbivory in a low Arctic wetland alters intraspecific plant root traits with consequences for carbon and nitrogen cycling. Journal of Ecology, 113:5, 1225-1238. doi: 10.1111/1365-2745.70028
• Saunders, T., Adkins, J., Atwood, T., Waring, B., Beard, K.H, (2025). Goose herbivory effects on early-stage litter decomposition in coastal Alaskan wetlands. Plant and Soil, doi: doi.org/10.1007/s11104-025-07383-w
• Petit Bon, M., Beard, K.H, Bråthen, K.A, Lee, H., Klöfkorn, R., Jónsdóttir, I.S, (2025). Goose grubbing and warming suppress summer net ecosystem CO2 uptake differentially across high-Arctic tundra habitats. Ecology, 106:1, e4498. doi: DOI: 10.1002/ecy.4498
• Smith, M.D, Wilkins, K.D, Holdrege, M.C, Wilfahrt, P., Collins, S.L, Knapp, A.K, Sala, O.E, Dukes, J.S, Phillips, R.P, Yahdjian, L., Gherardi, L., Ohlert, T., Beier, C., Fraser, L., Jentsch, A., Loik, M., Maestre, F., Power, S., Yu, Q., Felton, A., Munson, S., Luo, Y., Abdoli, H., Abedi, M., Alados, C., Alberti, J., Alon, M., An, H., Anacker, B., Anderson, M., Auge, H., Bachle, S., Bahalkeh, K., Bahn, M., Batbaatar, A., Bauerle, T., Beard, K.H, Behn, K., Beli, I., Biancari, L., Blindow , I., Bondaruk, V., Borer, E., Bork, E., Bruschetti, C., Byrne, K., Cahill, J., Calvo, D., Carbognani, M., Cardoni, A., Carlyle, C., Castillo-Garcia, M., Chang, S., Chieppa, J., Cianciaruso, M., Cohen, O., Cordiero, A., Cusack, D., Dahlke, S., Daleo, P., D'Antonio, C., Diettrich, L., Doherty, T., Dubbert, M., Ebeling, A., Eisenhauer, N., Fischer, F., Forte, T., Gebauer, T., Gozalo, B., Greenville, A., Guidoni-Martins, k., Hannusch, h., Haugum, S., Hautier, Y., Hefting, M., Henry, H., Hoss, D., Ingrisch, J., Iribarne, O., Isbell, F., Johnson, Y., Jordan, S., Kelly, E., Kimmel, K., Kreyling, J., Kröel-Dulay, G., Kröpfl, A., Kübert, A., Kulmatiski, A., Lamb, E., Larsen, K., Larsen, J., Lawson, J., Leder, C., Linstädter, A., Liu, J., Liu, S., Lodge, A., Longo, G., Loydi, A., Luan, J., Lubbe, F., Macfarlane, C., Mackie-Haas, K., Malyshev, A., Maturano-Ruiz, A., Merchant, T., Metcalfe, D., Mori, A., Mudongo, E., Newman, G., Nielsen, U., Nimmo, D., Niu, Y., Nobre, P., O'Connor, R., Ogaya, R., Oñatibia, G., Orbán, I., Osborne, B., Others, , (2024). Extreme drought impacts have been underestimated in grasslands and shrublands globally. Proceedings of the National Academy of Sciences, 121:4, e2309881120.
• Marchetti, J., French, S., Virgin, E.E, Lewis, E.L, Ki, K.C, Sermersheim, L.O, Brusch, G., Beard, K.H, (2024). Invading nonnative frogs use different microhabitats and change physiology along an elevation gradient. Journal of Experimental Zoology Part A: Ecological and Integrative Physiology
• Shi, B., Delgado-Baquerizo, M., Knapp, A., Smith, M., Reed, S., Osborne, B., Yolima , C., Maestre, F.T, Zhu, Y., Chen, A., Wilkins, K., Holdrege, M., Kulmatiski, A., Picon-Cochard, C., Roscher, ., Power, S., Byrne, .M, Churchill, A.C, Jentsch, A., Henry, H.A, Beard, K.H, Schuchardt, M.A, Eisenhauer, ., Otfinowski, ., Hautier, ., Shen, H., Wang, Y., Wang, Z., Wang, C., Cusack, D.F, Petraglia, A., Carbognani, M., Forte, T.G, Flory, S.L, Hou, P., Zhang, T., Gao, W., Sun, W., (2024). Aridity drives the response of soil total and particulate organic carbon to drought in temperate grasslands and shrublands. Science Advances, 10:40, doi: doi: 10.1126/sciadv.adq2654
• Petit Bon, M., Leffler, A.J, Kelsey, K.C, Williams, T.C, Beard, K.H, (2024). Projected near-future flooding and warming increase graminoid and plant-community biomass in high-latitude coastal wetland. Journal of Ecology, doi: doi:10.1111/1365-2745.14418
• Do Nascimento, L.A, Perez-Granados, C., Rodrigues Alencar, J.B, Beard, K.H, (2024). Time and habitat structure shape insect acoustic activity in the Amazon. Philosophical Transactions of the Royal Society B , 379:1904, 20230112 . doi: doi: 10.1098/rstb.2023.0112
• Kulmatiski, A., Holdrege, M.C, Chirvasa, C., Beard, K.H, (2024). Root distributions predict shrub-steppe responses to precipitation intensity. Biogeosciences, 21, 131–143. doi: 10.5194/bg-21-131-2024
• Marchetti, J., Beard, K.H, Virgin, E.E, Lewis, E.L, Hess, S.C, Ki, K.C, Sermersheim, L.O, Furtado, A.P, French, S., (2023). Invasive frogs show persistent physiological differences to elevation and acclimate to colder temperatures. Journal of Thermal Biology, 114, 103590.
• Holdrege, M.C, Kulmatiski, A., Beard, K.H, Palmquist, K., (2023). Precipitation Intensification Increases Shrub Dominance in Arid, Not Mesic, Ecosystems. Ecosystems, 26, 568–584. doi: https://doi.org/10.1007/s10021-022-00778-1
• Saunders, T., , Beard, K.H, Atwood, T., , (2023). Herbivores influence biogeochemical processes by altering litter quality and quantity in a subarctic wetland. Biogeochemistry, 166, 67-85. doi: 10.1007/s10533-023-01098-9
• Petit Bon, M., Hansen, B.B, Loonen, M.J, Petraglia, A., Bråthen, K., Böhner, H., Layton-Matthews, K., Beard, K.H, Le Moullec, M., Jónsdóttir, I.S, van der Wal, R., (2023). Long-term herbivore removal experiments reveal how geese and reindeer shape vegetation and ecosystem CO2-fluxes in high-Arctic tundra. Journal of Ecology
• Lacerda, J., Cafofo, E., Zocca, C., DeAraujo, A., Beard, K.H, Toledo, L., Ferreria, R., (2023). Plant-borne vibration is related to the vocal repertoire of an Atlantic forest Marsupial frog. Salamandra, 59:3, 275-283.
• Beard, K.H, Kelsey, K.C, Choi, R.T, Welker, J.M, Leffler, A.J, (2023). Goose Feces Effects on Subarctic Soil Nitrogen Availability and Greenhouse Gas Fluxes. Ecosystems, 26:1, 187-200.
• Kulmatiski, A., Beard, K.H, (2022). A modern two-layer hypothesis helps resolve the 'savanna problem'. Ecology Letters, 25:9, 1952-1960. doi: doi: 10.1111/ele.14067
• Holdrege, M., Kulmatiski, A., Beard, K.H, Palmquist, K., (2022). Precipitation intensification increases shrub growth in arid, not mesic, systems. Ecosystems, doi: doi: 10.1007/s10021-022-00778-1
• Choi, R., Petit Bon, M., Leffler, A.J, Kelsey, K.C, Welker, J.M, Beard, K.H, (2022). Short-term effects of experimental goose grazing and warming differ in three low-Arctic coastal wetland plant communities. Journal of Vegetation Science, 33:3, e13139 . doi: https://doi.org/10.1111/jvs.13139
• Cayuela, H., Lemaître , J., Léna, J., Ronget, V., Martínez-Solano, I., Clavel, J., Muths, E., Pilliod, D., Schmidt, B., Sánchez-Montes, G., Gutiérrez-Rodríguez, J., Pyke, G., Grossenbacher, K., J.B, Beard, K.H, Woolbright, L., (2022). Sex heterogamety drives the evolution of sex differences in aging rate across amphibians. Evolution, 76:2, 346-356. doi: doi: 10.1111/evo.14410
• Do Nascimento, L., Perez-Granados, C., Beard, K.H, (2021). Passive acoustic monitoring and automatic detection of the diel and acoustic structure of howler monkey roars. Diversity, 13, 566. doi: https:/doi.org/10.3390/d13110566
• Beard, K.H, Marchetti, J.R, (2021). Predator-prey reunion: Non-native coqui frogs avoid their native predators. Ichthyology and Herpetology, 109:3, 791-795.
• Holdrege, M., Beard, K.H, Kulmatiski, A., (2021). Winter wheat growth resistant to increases in rain and snow intensity. Agronomy, 11:4, 751. doi: 10.3390/agronomy11040751
• Foley, K., Beard, K.H, Atwood, T., Waring, B., (2021). Herbivory changes soil microbial communities and greenhouse gas fluxes in high-latitude wetlands. Microbial Ecology, doi: https://doi.org/10.1007/s00248-021-01733-8
• Holdrege, M.C, Beard, K.H, Kulmatiski, A., (2021). Woody plant growth increases with precipitation intensity in a cold semi-arid system. Ecology, 102:1, e03212 . doi: doi: 10.1002/ecy.3212
• DoNascimento, L., Campos-Cerqueira, M., Beard, K.H, (2020). Acoustic metrics predict habitat type and vegetation structure in the Amazon. Ecological Indicators, 117, 106679. doi: 10.1016/j.ecolind.2020.106679
• Beard, K.H, Durham, S., Willig, M., Zimmerman, J., (2020). Lizard and frog removal increases spider abundance but does not cascade to increase herbivory. Biotropica
• Kulmatiski, A., Beard, K.H, Holdrege, M.C, February, E., (2020). Small differences in root distributions allow resource niche partitioning. Ecology and Evolution, 10:18, 9776-9787. doi: 10.22541/au.159056294.45813944
• Atwood, T., Valentine, S.A, Hammill, E., McCauley, D.J, Madin, E.M, Beard, K.H, Pearse, W., (2020). Herbivores at the highest risk of extinction among mammals, birds, and reptiles. Science Advances, 6:32, doi: 10.1126/sciadv.abb8458
• Choi, R.T, Beard, K.H, Kelsey, K., Leffler, A.J, Schmutz, J., Welker, J., (2020). Early goose arrival increases soil nitrogen availability more than an advancing spring in coastal western Alaska. Ecosystems, 23, 1309-1324. doi: doi: 10.1007/s10021-019-00472-9
• Lourenco-de-Moraes, R., Campos, F., Ferreira, R.B, Beard, K.H, Sole, M., Llorente, G., Bastos, R., (2020). Functional traits explain amphibian distribution in the Brazilian Atlantic Forest. Journal of Biogeography, 47:1, 275-287.
• Lantyer-Silva, A., Lirio, F., Tonini, J., Zocca, C., Fraga, C., Waichert, C., E.K, Marciano-Jr, ., Lacerda, J., Alves, J., Beard, K.H, Cipriano, R., Ferreira, R., (2019). Projecto bromeligenous: Aliando pesquisa e educação em prol da conservação de anfíbios de bromélia. Herpetologia Brasileira, 8:3, 20-28.
• Beard, K.H, Kelsey, K., Leffler, A.J, Welker, J., (2019). The missing angle: Ecosystem consequences of phenological mismatch. Trends in Ecology and Evolution, 34:10, 19-22. doi: 10.1016/j.tree2019.07.019
• Leffler, A.J, Beard, K.H, Kelsey, K., Choi, R.T, Schmutz, J., Welker, J., (2019). Cloud cover and delayed herbivory relative to timing of spring onset interact to dampen climate change impacts on net ecosystem exchange in a coastal Alaskan wetland. Environmental Research Letters, 14:8, 084030. doi: 10.1088/1748-9326/ab1c91
• Ferreira, R.B, Monico, A.T, Zocca, C.Z, Santo, M.T, Lirio, F.C, Tonini, J.F, Sabagh, L.M, Cipiano, R., Waichert, C., Crump, M., Beard, K.H, Toledo, L., Duca, C., (2019). Uncovering the natural history of the bromeligenous frog, Crossodactylodes izecksohi (Leptodactlylidae, Paratelmatobiinae). South-American Journal of Herpetology, 14:2, 136-145. doi: 10.2994/SAJH-D-17-00092.1
• Ferreira, R.B, Monico, A.T, Silva, E.T, Lirio, F.C, Zocca, C.Z, Mageski, M.M, Tonini, J.F, Beard, K.H, Duca, C., Silva-Soares, T., (2019). Amphibians of Santa Teresa, Brazil: the hotspot further evaluated. Zookeys, 857, 139-162. doi: 10.3897/zookeys.857.30302
• Ferreira, R.B, Lourenço-de-Moraes, R., Zocca, C., Duca, C., Beard, K.H, Brodie, E.D, (2019). Antipredator mechanisms of post-metamorphic anurans: a global database and classification system. Behavior Ecology and Sociobiology, 73, 69. doi: 10.1007/s00265-019-2680-1
• Hill, S.A, Beard, K.H, S.S, A.S, (2019). Invasive coqui frogs are associated with differences in small Indian mongoose and rat abundances and diets in Hawaii. Biological Invasions, 21:6, 2177-2190. doi: 10.1007/s10530-019-01965-3
• Lourenço-de-Moraes, M., Camp, F., Ferreira, R., Sole, M., Beard, K.H, Bastos, R., (2019). Back to the future: Conserving functional and phylogenetic diversity in the amphibian climate refuges. Biodiversity and Conservation, 28:5, 1049-1073. doi: 10.1007/s10531-019-01706-x
• Kulmatiski, A., Beard, K.H, (2019). Chronosequence and direct observation approaches reveal complementary community dynamics in a novel ecosystem. Plos One, 14:3, e0207047. doi: 10.1371/journal.pone.0207047
• Beard, K.H, Choi, R.T, Leffler, A.J, Carlson, L.G, Kelsey, K., Schmutz, J., Welker, J., (2019). Migratory goose arrival time plays a larger role in influencing forage quality than advancing springs in an Arctic coastal wetland. PlosOne, 14:3, e0213037. doi: 10.1371/journal.pone.0213037
• Kalnicky, E., Brunson, M.W, Beard, K.H, (2019). Predictors of Participation in Invasive Species Control Activities Depend on Prior Experience with the Species. Environmental Management, 63:1, 60-68. doi: 10.1007/s00267-018-1126-2
• Costello, M., Beard, K.H, Primack, R., DeVictor, V., Bates, A., (2019). Editorial: Are killer bees good for coffee? Can I increase citations of my paper through framing its title? . Biological Conservation, 229, A1-A5. doi: 10.1016/j.biocon.2018.07.010
• Leffler, A.J, Beard, K.H, Kelsey, K., Choi, R.T, Schmutz, J., Welker, J., (2019). Delayed grazing by migratory geese increases summer-long CO2 uptake in coastal Western Alaska . Global Change Biology, 25:1, 277-289. doi: 10.1111/gcb.14473
• Henry, H., Abedi, M., Alados, C., Beard, K.H, Fraser, L., Jentsch, , Kreyling, , Kulmatiski, A., Lamb, J., Sun, W., Vankoughnett, M., Venn, S., Werner, C., Beil, I., Blindow, I., Dahlke, S., Dubbert, M., Effinger, A., Garris, H., Gartzia, M., Gebauer, T., Arfin Khan, M., Malyshev, A., Paulson, J., Pueyo, Y., Stover , ., Yang, X., (2018). Increased soil frost versus summer drought as drivers of plant biomass responses to reduced precipitation: results from a globally-coordinated field experiment. Ecosystems, 21:7, 1432-1444. doi: 10.1007/s10021-018-0231-7
• Koerner, S., Smith, ., Burkepile, D., Hanan, N., Avolio, M., Collins, S., Knapp, A., Lemoine, N., Beard, K.H, Kulmatiski, A., Veblen, K.E, (2018). Change in dominance determines herbivore effects on plant biodiversity. Nature Ecology and Evolution, 2:12, 1925-1932.
• Adler, P., Smull, D., Beard, K.H, Choi, R.T, Furniss, T., Kulmatiski, A., Meiners, J., Tredennick, A.T, Veblen, K.E, (2018). Competition and coexistence in plant communities: intraspecific competition is stronger than interspecific competition. Ecology Letters, 21:9, 1319–1329. doi: 10.1111/ele.13098
• Kelsey, K., Leffler, A.J, Beard, K.H, Choi, R.T, Schmutz, J., Welker, J., (2018). Developing phenological mismatch in coastal western Alaska may increase summer season greenhouse gas uptake. Environmental Research Letters, 13:4, 044032. doi: 10.1088/1748- 9326/aab698
• Carlson, L.G, Beard, K.H, Adler, P., (2018). Direct effects of warming increase woody plant abundance in a subarctic wetland. Ecology and Evolution, 8:5, 2868–2879. doi: 10.1002/ece3.3902
• O'Neill, E.M, Beard, K.H, Fox, C.W, (2018). Life history traits in native and introduced populations of the coqui frog . Copeia, 106:1, 161-170. doi: 10.1643/CE-17-642
• Smith, R.L, Koons, D.N, Beard, K.H, (2018). Invasive coqui frogs are associated with greater abundances of non-native birds in Hawaii. The Condor: Ornithological Applications, 120:1, 16-29.
• Kulmatiski, A., Beard, K.H, Norton, J.M, Heavilin, J.E, Forero, L., Grezer, J., (2017). Live long and prosper: plant-soil feedback, lifespan and landscape abundance covary. Ecology, 98:12, 3063-3073. doi: 10.1002/ecy.2011
• Kulmatiski, A., Sprouse, S., Beard, K.H, (2017). Soil type more than precipitation determines fine-root abundance in savannas of Kruger National Park, South Africa. Plant and Soil, 417:1-2, 523-333. doi: 10.1007/s11104-017-3277-y
• Smith, R.L, Beard, K.H, Shiels, A.B, (2017). Different prey resources suggest little competition between non-native frogs and insectivorous birds despite isotopic niche overlap. Biological Invasions, 19:3, 1001-1013. doi: 10.1007/s10530-016-1333-9
• Marsh, D.M, Cosentino, B.J, Jones, K.S, Apodaca, J.J, Beard, K.H, Bell, J.M, Bozarth, C., Carper, D., Charbonnier, J.F, Dantas, A., Forys, E.A, Foster, M., General, J., Genet, K.S, Hanneken, M., Hess, K.R, Hill, S.A, Karraker, N. , Kilpatrick , E., Langen, T.A, Langofrd, J., Lauer, K., McCarthy, A.J, Neale, J., Patel, S., Patton, A., Soutwick, C., Stearrett, N., Steijn, N., Tasleem, M., Taylor, J.M, Vonesh, J.R, (2017). Effects of road and land use on pond-breeding amphibians: patterns across spatial scales and regions in the eastern and central United States. Diversity and Distributions, 23:2, 158-170.
• Kelsey, K.C, Leffler, A.J, Beard, K.H, Schmutz, J.A, Choi, R.T, Welker, J.M, (2016). Interactions among climate, topography and herbivory control greenhouse gas (CO2, CH4, and N2O) fluxes in a subarctic coastal wetland. Journal of Geophysical Research - Biogeosciences, 121:12, 2960–2975. doi: 10.1002/2016JG003546
• Tonini, J.F, Beard, K.H, Ferreira, R.B, Pyron, R.A, (2016). Fully-sampled phylogenies for squamates reveal evolutionary patterns in extinction risk. Biological Conservation, 204, 22-31.
• Wallis, A.C, Smith, R.L, Beard, K.H, (2016). Temporal foraging patterns of non-native Coqui frogs (Eleutherodactylus coqui) in Hawaii. Journal of Herpetology, 50:4, 582-588.
• Ferreira, R.B, Lourenço-de-Moraes, R., Teixeira, ,.L, Beard, K.H, (2016). Frogs associations with bromeliads in an abandoned cacao plantation in northeastern Brazil. Northwestern Journal of Zoology, 12:2, 392-396. doi: e162501
• Costello, M.J, Beard, K.H, Corlett, R., Cumming, G., Loyola, R., Maas, B., Miller-Rushing, A., Pakeman, R., Primack, R., (2016). Field work ethics in biological research: Viewpoint of Biological Conservation editors. Biological Conservation, 203, 268-271. *
• Kulmatiski, A., Beard, K.H, Grezer, J., Forero, L., Heavilin, J.E, (2016). Using plant-soil feedbacks to predict plant biomass in diverse communities. Ecology, 97:8, 2064-2073. doi: 10.1890/15-2037.1
• Ferreira, R.B, Beard, K.H, Crump, M., (2016). Breeding guilds determine frog response to edge effects in Brazil’s Atlantic Forest. . PLOS One, 11:6, e0156781.
• Tingley, R., Mahoney, P.J, Durso, A.M, Taillan, A.G, Morán-Ordóñez, A., Beard, K.H, (2016). Threatened and invasive reptiles are not two sides of the same coin. Global Ecology and Biogeography, 25:9, 1050–1060 .
• Mageski, M.M, Ferreira, R.B, Beard, K.H, Costa, L.C, Jesus, P.R, Medeiros, C.C, Ferreira, P.D, (2016). Bromeliad selection by Phyllodytes luteolus (Anura, Hylidae): the influence of plant structure and water quality factors . Journal of Herpetology, 50:1, 108-112.
• Ferreira, R.B, Faivovich, J., Beard, K.H, Pombal Jr., J., (2015). The First Bromeligenous Species of Dendropsophus (Anura: Hylidae) from Brazil's Atlantic Forest. PLOS One, doi: 10.1371/journal.pone.0142893
• Warren, C., Kulmatiski, A., Beard, K.H, (2015). A combined tracer/evapotranspiration model approach estimates plant water uptake in native and non-native shrub-steppe communities. Journal of Arid Environments, 121:2015, 67–78.
• Ferreira, R.B, Beard, K.H, Choi, R.T, Pitt, W.C, (2015). Diet of the non-native Greenhouse Frog (Eleutherodactylus planirostris) in Maui, Hawaii. Journal of Herpetology, 49:4, 586-593.
• Nolan, N.E, Kulmatiski, A., Beard, K.H, Norton, J.M, (2015). Activated carbon decreases invasive plant growth by mediating plant-microbe. AoB Plants, 7, plu072. doi: doi: 10.1093/aobpla/plu072
• Mahoney, P., Beard, K.H, Durso, A.M, Taillan, A.G, Long, A.L, Kindermann, R.J, Nolan, N.E, Kinka, D., Mohn, H.E, (2015). Introduction effort, climate matching, and species traits as predictors of global establishment success in non-native reptiles. Diversity and Distributions, 21:1, 64-74. doi: 10.1111/ddi.12240
• Cosentino, B.J, Marsh, D.M, Jones, K.S, Apodaca, J.J, Bates, C., Beach, J., Beard, K.H, Becklin, K., Bell, J., Crockett, C., Fawson, G., Fjelsted, J., Forys, E.A, Genet, K.S, Grover, M., Holmes, J., Indeck, K., Karraker, N. , Kilpatrick , E., Langen, T.A, Mugel, S.G, Molina, A., Vonesh, J.R, Weaver, R., Willey, A., (2014). Citizen science reveals widespread negative effects of roads on amphibian distributions. Biological Conservation, 180, 31–38. doi: 10.1016/j.biocon.2014.09.027
• Kalnicky, E.A, Brunson, M.W, Beard, K.H, (2014). A social-ecological systems approach to non-native species: Habituation and its impact on management of coqui frogs in Hawaii. Biological Conservation, 180, 187–195. doi: 10.1016/j.biocon.2014.09.044
• Kulmatiski, A., Anderson-Smith, A., Beard, K.H, Doucette-Riis, S., Mazzacavallo, M., Nolan, N.E, Ramirez, R., Stevens, J.R, (2014). Most soil trophic guilds increase plant growth: a meta-analytical review. Oikos, 123:2, 1409-1419. doi: 10.1111/oik.01767
• Olson, C.A, Diesmos, A.C, Beard, K.H, (2014). Geographical Distribution: Eleutherodactylus planirostris (greenhouse frog). , 45:4, 652-653.
• Kalnicky, E.A, Beard, K.H, Brunson, M.W, (2013). Community-level response to habitat structure manipulations: An experimental case study in a tropical ecosystem. Forest Ecology and Management, 307, 313-321.
• Beard, K.H, Faulhaber, C.A, Howe, F.P, Edwards, T.C, (2013). Rodent-mediated interactions among seed species of differing quality in a shrubsteppe ecosystem. Western North American Naturalist, 73:4, 426–441.
• Kalnicky, E.A, Beard, K.H, Brunson, M.W, (2013). Community-level response to habitat structure manipulations: An experimental case study in a tropical ecosystem. Forest Ecology and Management, 307, 313-321.
• Kulmatiski, A., Beard, K.H, (2013). Increasing precipitation intensity linked to woody plant encroachment. Nature Climate Change, 3, 833-837.
• Hoopes, M., Marsh, D., Beard, K.H, Goldberg, N., Aparicio, A., Hixon, B., Laflower, D., Lee, L., Little, A., Mooney, E., Pallette, A., Ravenscraft, A., Scheele, S., Stowe, K., Skyes, C., Watson, R., Yang, B., (2013). Invasive plants in U.S. National Wildlife Refuges: A coordinated research project using undergraduate ecology students. Bioscience, 63:8, 644-656.
• Poessel, S., Beard, K.H, Callahan, C., Rodrigo, F., (2013). Biotic acceptance in introduced amphibians and reptiles in Europe and North America. Global Ecology and Biogeography, 22, 192–201. doi: DOI: 10.1111/j.1466-8238.2012.00797.x
• Kulmatiski, A., Beard, K.H, (2013). Root niche partitioning among grasses, saplings, and trees measured using a tracer technique. Oecologia, 171:1, 25-37. doi: 10.1007/s00442-012-2390-0
• Ferreira, R.B, Beard, K.H, Peterson, S.L, Poessel, S.A, Callahan, C.M, (2012). Establishment of introduced reptiles increases with the presence and richness of native congeners. Amphibia-Reptilia, 33:2012, 387-392. doi: 10.1163/15685381-00002841
• Ferreira, R.B, Callahan, C.M, Poessel, S.A, Beard, K.H, (2012). Global assessment of establishment success for amphibian and reptile invaders. Wildlife Research, 39:7, 637-640. doi: DOI: 10.1071/WR12051
• O'Neill, E.M, Beard, K.H, Pfrender, M.E, (2012). Cast adrift on an island: introduced populations experience an altered balance between selection and drift. Biology Letters, 8:5, 890-893. doi: 10.1098/rsbl.2012.0312
• Pitt, W.C, Beard, K.H, Doratt, R.E, (2012). Management of invasive coqui frog populations in Hawaii. Outlooks on Pest Management, 23:4, 166-169 . doi: 10.1564/23jun00
• Kulmatiski, A., Beard, K.H, Heavilin, J.E, (2012). Plant-soil feedbacks provide an alternative explanation for diversity-productivity relationships. Proceedings of the Royal Society B, 279:1740, 3020–3026. doi: 10.1098/rspb.2012.0285
• Olson, C.A, Beard, K.H, Pitt, W.C, (2012). Biology and Impacts of Pacific Island Invasive Species: Eleutherodactylus planirostris, the Greenhouse frog (Anura: Leptodactylidae). Pacific Science, 66:3, 255-270.
• Bisrat, S.A, White, M.A, Beard, K.H, Cutler, D., (2012). Predicting the distribution potential of an invasive Puerto Rican frog (Eleutherodactylus coqui) in Hawaii using remote sensing data. Diversity and Distributions, 18:7, 648-660. doi: 10.1111/j.1472-4642.2011.00867.x
• Choi, R., Beard, K.H, (2012). Coqui frog invasions change invertebrate communities in Hawaii. Biological Invasions, 14:5, 939-948. doi: 10.1007/s10530-011-0127-3
• Olson, C.A, Beard, K.H, Koons, D.N, Pitt, W.C, (2012). Detection probability of two introduced frogs in Hawaii: implications for assessing non-native species distributions. Biological Invasions, 14:4, 889-900 . doi: 10.1007/s10530-011-0125-5
• Olson, C.A, Beard, K.H, (2012). Diet of the invasive greenhouse frog in Hawaii. Copeia, 2012:1, 122-130. doi: DOI: 10.1643/CE-11-008
• Kulmatiski, A., Beard, K.H, (2011). Changing soils to manage plant communities: Activated carbon as a restoration tool in ex-arable fields. Restoration Ecology, 19:101, 102-110.
• O'Neill, E.M, Beard, K.H, (2011). Clinal variation in calls of native and introduced populations of Eleutherodactylus coqui. Copeia, 2011:1, 18–28.
• Kulmatiski, A., Heavilin, J.E, Beard, K.H, (2011). Testing predictions of a three-species plant-soil feedback model. Journal of Ecology, 99:2, 542-550. doi: doi:10.1111/j.1365- 2745.2010.01784.x
• Gilbert-Norton, L., Wilson, R., Stevens, J.M, Beard, K.H, (2010). Corridors increase movement: A meta-analytical review.. Conservation Biology, 24:3, 660-668.
• Kulmatiski, A., Beard, K.H, Meyerson, L.A, Gibson, J.C, Mock, K.E, (2010). Nonnative Phragmites australis invasion into Utah wetlands. Western North American Naturalist, 70, 541-552.
• Kulmatiski, A., Beard, K.H, Verweij, R.J, February, E.C, (2010). A depth-controlled tracer technique quantifies the location, extent, and timing of water uptake in a sub-tropical savanna. New Phytologist, 188:1, 199-209.
• Tuttle, N.C, Beard, K.H, Pitt, W.C, (2009). Invasive litter, not an invasive insectivore, determines invertebrate communities in Hawaiian forests. Biological Invasions, 11:4, 845–855.
• Peacock, M.M, Beard, K.H, O'Neill, E.M, Kirchoff, V., Peters, M.B, (2009). Strong founder effects and low genetic diversity in introduced populations of Coqui frogs. Molecular Ecology, 18, 3603-3615.
• Beard, K.H, Price, E.A, Pitt, W.C, (2009). Biology and impacts of Pacific Island invasive species: Eleutherodactylus coqui, the Coqui frog (Anura: Leptodactylidae). Pacific Science, 63(3), 297-316.
• Peters, M.P, Beard, K.H, Hagen, C., O'Neill, E.M, Mock, K.E, Pitt, W.C, Glenn, T.C, (2008). Isolation of microsatellite loci from the coqui frog, Eleutherodactylus coqui. Molecular Ecology Notes8, 139-141. doi: 10.1111/j.1471-8286.2007.01899.x
• Tuttle, N.C, Beard, K.H, Al-Chokhachy, R., (2008). Aerially applied citric acid reduces an invasive frog. Wildlife Research, 35:7, 676-683.
• Kulmatiski, A., Beard, K.H, Stevens, J.R, Cobbold, S.M, (2008). Plant-soil feedbacks: A meta-analytical review. Ecology Letters, 11:9, 980-992.
• Kyle, G.P, Kulmatiski, A., Beard, K.H, (2008). Pocket gophers change plant species composition in a shrub steppe ecosystem. Western North American Naturalist, 68:3, 374-381.
• Sin, H., Beard, K.H, Pitt, W.C, (2008). An invasive frog, Eleutherodactylus coqui, increases new leaf production and leaf litter decomposition rates through nutrient cycling in Hawaii. Biological Invasions, 10:3, 335-345.
• Bolger, D.T, Beard, K.H, Suarez, A., Case, T., (2008). Increased abundance of native and non-native spiders with habitat fragmentation. Diversity and Distributions, 14:4, 655-665.
• Beard, K.H, Al-Chokhachy, R., Tuttle, N.C, O'Neill, E.M, (2008). Population density estimates and growth rates of Eleutherodactylus coqui in Hawaii. Journal of Herpetology, 42:4, 626-636.
• Kulmatiski, A., Beard, K.H, (2008). Decoupling plant-growth from land-use legacies in soil microbial communities.. Soil Biology and Biogeochemistry, 40(5), 1059-1068.
• Cutler, D., Edwards, T.C, Beard, K.H, Cutler, A., Hess, K.T, Gibson, J., Lawler, J.J, (2007). Random Forests for Classification in Ecology. Ecology, 88:11, 2783-2792.
• Velo-Antón, G., Burrowes, P.A, Joglar, R., Martínez-Solano, I., Parra-Olea, G., Beard, K.H, (2007). Phylogenetic study of Eleutherodactylus coqui (Anura: Leptodactylidae) reveals deep genetic fragmentation in Puerto Rico and origins in Hawaii. Molecular Phylogenetics and Evolution, 45, 716-728.
• Kyle, G.P, Beard, K.H, Kulmatiski, A., (2007). Reduced soil compaction enhances establishment of non-native plant species. Plant Ecology, 193(2), 223-232.
• Beard, K.H, (2007). Diet of the invasive frog, Eleutherodactylus coqui, in Hawaii.. Copeia, 2, 281-291.
• Rexroad, E.A, Beard, K.H, Kulmatiski, A., (2007). Vegetation responses to 35 and 55 years of native ungulate grazing in shrub steppe communities. Western North American Naturalist, 67(1), 16-25.
• Kulmatiski, A., Beard, K.H, Stark, J.M, (2006). Exotic communities shift water-use timing. Plant and Soil, 288:1-2, 271-284.
• Beard, K.H, Pitt, W.C, (2006). Potential predators of an invasive frog (Eleutherodactylus coqui) in Hawaiian forests. Journal of Tropical Ecology, 22:4, 1-3.
• Kulmatiski, A., Beard, K.H, (2006). Activated Carbon as a Restoration Tool. Restoration Ecology, 14:2, 251-257.
• Kulmatiski, A., Beard, K.H, Stark, J.M, (2006). Exotic plant communities shift water-use timing in a shrub-steppe ecosystem. Plant & Soil, 288, 271-284.
• Kulmatiski, A., Beard, K.H, Stark, J.M, (2006). Soil history as a primary control on plant invasion in abandoned agricultural fields. J Applied Ecology, 43, 868-876.
• Beard, K.H, Vogt, K.A, Vogt, D.J, Scatena, F.N, Covich, A., Siccama, T.C, Sigurdardottir, R., Crowl, T.A, (2005). Structural and functional responses of a subtropical forest to 10 years of hurricanes and droughts. Ecological Monographs, 75:3, 345-361.
• Beard, K.H, Pitt, W.C, (2005). Potential consequences of the coqui frog invasion in Hawaii. Diversity and Distributions, 11(5), 427-433.
• Beard, K.H, O'Neill, E.M, (2005). Infection of an invasive frog Eleutherodactylus coqui by the chytrid fungus Batrachochytrium dendrobatidis in Hawaii. Biological Conservation, 126(4), 591-595.
• Beard, K.H, Wang, D., Waite, C.E, Decker, K.L, Hawley, G.J, DeHayes, D.D, Hughes, J.W, Cumming, J.R, (2005). Quantifying the importance of abiotic and biotic factors in controlling nutrient export from a developing ecosystem. Ecosystems, 8, 210-224.
• Beard, K.H, (2005). Structural, functional characteristics and resilience of forest ecosystems impacted by multiple, severe disturbances in legacy environments. Ecological Monographs, 75(3), 345-361.
• Beard, K.H, (2004). Finding endemic soil-based controls for weed growth. Weed Technology, 18, 1353-1358.
• Kulmatiski, A., Beard, K.H, Stark, J.M, (2004). Finding endemic soil-based controls on weed growth. Weed Technology, 18, 115-120.
• Kulmatiski, A., Beard, K.H, (2004). Reducing sampler error in soil research. Soil Biology and Biochemistry, 36 (2), 383-385.
• Beard, K.H, Eschtruth, A.K, Vogt, K.A, Vogt, D.J, Scatena, F.N, (2003). Amphibian effects on its prey and ecosystem: Evidence from two spatial scales. Journal of Tropical Ecology, 37:19, 607-617.
• Kulmatiski, A., Vogt, D.J, Siccama, T.G, Beard, K.H, (2003). Detecting nutrient pool changes in rocky forest soils. Soil Science Society of America Journal, 67, 1282-1286.
• Beard, K.H, McCullough, S., Eschtruth, A.K, (2003). A quantitative assessment of habitat preferences for the Puerto Rican terrestrial frog, Eleutherodactylus coqui. Journal of Herpetology, 37(1), 10-17.
• Beard, K.H, (2003). The effects of the frog Eleutherodactylus coqui on invertebrates and ecosystem processes at two scales in the Luquillo Experimental Forest, Puerto Rico. Journal of Tropical Ecology, 19, 607-617.
• Vogt, K.A, Beard, K.H, Hammann, S., O'Hara, J.L, Vogt, D.J, Scatena, F.N, Parry, B.A, (2002). Indigenous Knowledge Informing Management of Tropical Forests: The Link between Rhythms in Plant Secondary Chemistry and Lunar Cycles. Ambio, 35:485-490
• Beard, K.H, Vogt, K.A, Kulmatiski, A., (2002). Top-down effects of a terrestrial frog on nutrient concentrations in a subtropical forest. Oecologia, 133, 583-593.
• Shelly, D.K, Beard, K.H, Hengarten, N.J, (2000). Animal-Distribution Modeling in Gap Analysis: An Evolving Science. Conservation Biology, 14(5), 1224-1225.
• Kiesecker, J.M, Skelly, D.K, Beard, K.H, Presser, E., (1999). Behavioral Reduction of Infection Risk in a Larval Amphibian. Proceedings., National Academy of Sciences of the United States of America, 96(16), 9165-9168.
• Beard, K.H, Hengarten, N., Skelly, K., (1999). The effectiveness of predicting breeding bird distributions using probabilistic models. Conservation Biology, 13(5), 1108-1116.
• Beard, K.H, DePriest, P.T, (1996). Genetic variation within and among mats of the reindeer-lichen, Cladina subtenuis (des abb.). Hale & W. Culb. Lichenologist, 28(2), 171-182.
• Choi, R.T, Beard, K.H, Leffler, A.J, Kelsey, K., Schmutz, J., Welker, J.M, Phenological mismatch between season advancement and herbivory timing alters Arctic plant traits. Journal of Ecology, doi: 10.1111/1365-2745.13191

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